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To attach a file to the record: an invoice, a photo, a contract.
How formulas and the API refer to this field. It can't change once the page is saved.
The server enforces a hard ceiling of 50 MB regardless of this setting.
On this page the file isn't uploaded anywhere: you can pick one and download it, but it isn't kept. In an application, the file is uploaded and stays with the record.
Each file keeps its versions: uploading a new one in place of the old one doesn't lose the old one. Who may download files depends on their access to the application.